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| Subspecies: | Unknown |
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| Est. World Population: | |
| CITES Status: | NOT LISTED |
| IUCN Status: | Near Threatened |
| U.S. ESA Status: | NOT LISTED |
| Body Length: | |
| Tail Length: | |
| Shoulder Height: | |
| Weight: | |
| Top Speed: | |
| Jumping Ability: | (Horizontal) |
| Life Span: | in the Wild |
| Life Span: | in Captivity |
| Sexual Maturity: | (Females) |
| Sexual Maturity: | (Males) |
| Litter Size: | |
| Gestation Period: | |
Habitat:
Roosts mainly in tree cavities, but occasionally uses slits under elevated bridges, buildings and bird boxes (Fukui 2015). Environments around roosts vary from forest to urban areas. Feeding habitats are unknown.
It is nocturnal but often shows day-time feeding mainly in early winter. It mainly feeds on Coleoptera, Lepidoptera, Trichoptera, Ephemeroptera and Hemiptera (Fukui et al. 2013). In spring and from autumn to early winter, this species regularly preys on small birds of Passeriformes (Fukui et al. 2013).
From June to August, females form maternity colonies (Maeda 1974). The size of a maternity colony varies from tens to more than a hundred individuals. During the period when females are involved in maternity colonies, males roost in solitary or males form small colonies consisting of several to tens of individuals (Maeda 1974). Parturition occurs during late June to early July in Iwate and Hokkaido (Maeda 1972). Litter size is sometimes 1 but usually 2. Forearm length at birth and its ratio to mother are 22.1 mm (range = 20.0–27.0; n = 10) and 0.36 (n = 10) on average respectively; body weight at birth is 6.3 g on average (5.4–8.3; n = 9), and increases to 25 g up to 4 weeks of age; young begin to fly at about 40–45 days of age (Maeda 1972). Yearling females copulate in the first autumn but yearling males do not; the testes of adult males reach a maximum size in July and this state continues until the beginning of October (Maeda 1974).
It is nocturnal but often shows day-time feeding mainly in early winter. It mainly feeds on Coleoptera, Lepidoptera, Trichoptera, Ephemeroptera and Hemiptera (Fukui et al. 2013). In spring and from autumn to early winter, this species regularly preys on small birds of Passeriformes (Fukui et al. 2013).
From June to August, females form maternity colonies (Maeda 1974). The size of a maternity colony varies from tens to more than a hundred individuals. During the period when females are involved in maternity colonies, males roost in solitary or males form small colonies consisting of several to tens of individuals (Maeda 1974). Parturition occurs during late June to early July in Iwate and Hokkaido (Maeda 1972). Litter size is sometimes 1 but usually 2. Forearm length at birth and its ratio to mother are 22.1 mm (range = 20.0–27.0; n = 10) and 0.36 (n = 10) on average respectively; body weight at birth is 6.3 g on average (5.4–8.3; n = 9), and increases to 25 g up to 4 weeks of age; young begin to fly at about 40–45 days of age (Maeda 1972). Yearling females copulate in the first autumn but yearling males do not; the testes of adult males reach a maximum size in July and this state continues until the beginning of October (Maeda 1974).
Range:
Palaearctic, marginally Oriental. This species is known from eastern China (provinces of Heilongjiang, Jilin, Henan, Zhejiang, and Anhui (Smith and Xie 2013), the Korean Peninsula (Yoon 2010) and Japan, where there are records from Hokkaido, Honshu, Shikoku, Kyushu and Isls of Iki, Fukue, Okinawa-jima and Tsushima (Fukui 2015). In the Russian Far East, it is known from several doubtful observations, there are no significant evidences for presence of this bat in Russian fauna (Tiunov 1997), however, there is high possibility for its occurrence at least as vagrant.
Conservation:
It is listed as Near Threatened (NT) in the Chinese Red List, Vulnerable (VU) in the Japanese Red List (Ministry of the Environment 2014), and as a Rare Species in Russia.




